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| Upper Huallaga River Valley, Peru; 5/96 ssp signatus (formerly considered a distinct species, E harmodius) | female same data |
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| Napo River Valley 1100 meters Ecuador ssp xeniades |
Transandean from Colombia to Bolivia at moderate elevations. Several subspecies:
isus - W Andes valleys
in Colombia, rare
halex - E andes in
Colombia
xeniades - E Ecuador
tabaconas - S Ecuador
and N Peru
imaus - Huallaga
Valley of Peru
signatus - Central
Peru to Bolivia