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| Tingo Maria, Peru; 5/95 ssp. serville | Rio San Juan, Colombia 1600 meters April 1977 ssp colombus |
Transandean species found from Columbia south to Bolivia generally at moderate elevations in Andean mountain valleys. Ssp colombus has been considered by some to represent a full species, and I tend to support this notion, though I have kept it here to emphasize their relationship to each other
colombus - Central and western Colombia in
valleys of Pacific flowing rivers
acritus - Columbia and W Venezuela in Eastern
Andes
serville - common from S Colombia to S Bolivia
in Andean river valleys